Below is an excerpt from Ng’weno et al. 2019 (Frontiers in E…

Below is an excerpt from Ng’weno et al. 2019 (Frontiers in Ecology & Evolution): “Zebra populations are controlled bottom-up by rainfall, are the primary prey of lions, and support a growing population of lions since the cessation of lethal control in the late-1980s. Although they are less abundant than zebra, hartebeest are selectively killed by lions and are controlled top-down, such that hartebeest incur negative, indirect impacts from zebra via lions.” What is the best description of this type of competition?